{"id":9379,"date":"2021-09-20T20:30:44","date_gmt":"2021-09-21T00:30:44","guid":{"rendered":"https:\/\/www.bumc.bu.edu\/anatneuro\/?page_id=9379"},"modified":"2025-07-31T09:39:32","modified_gmt":"2025-07-31T13:39:32","slug":"phd-program","status":"publish","type":"page","link":"https:\/\/www.bumc.bu.edu\/anatneuro\/phd-program\/","title":{"rendered":"PhD Program"},"content":{"rendered":"<\/div>\n<p><!-- ___________________Navbar___________________ --><\/p>\n<div id=\"navvy\"><\/div>\n<p><script type=\"text\/javascript\" src=\"\/anatneuro\/files\/2025\/05\/nav-5-2025.js\"><\/script><\/p>\n<div class=\"mobile-title mobile-banner\">\n<p class=\"mobile-header\">PhD in Anatomy &#038; Neurobiology<\/p>\n<\/div>\n<style>.mobile-banner{background-image: url(\"\/anatneuro\/files\/2019\/08\/dlx5_6cre-lnl-gfp-e14-to-p8-montage-dtbr-2-1.jpg\"); background-size: cover;}<\/style>\n<p><!-- ___________________Banner___________________ --><\/p>\n<div class=\"phd-parallax\">\n<div class=\"true-nav nav-border\"><\/div>\n<div class=\"title-padding\">\n<p class=\"title-header\">PhD in Anatomy &#038; Neurobiology<\/p>\n<\/div>\n<\/div>\n<p><!-- ___________________Text___________________ --><\/p>\n<div style=\"padding-right: 50px; padding-left: 50px; padding-bottom: 20px;\">\n<p style=\"text-align: justify;\">This 64 credit doctoral program includes courses in Anatomy and Neuroscience that are offered within this department or are offered by related departments on the Medical Campus. An integral part of the program is the requirement for students to participate as Teaching Fellows in Medical and Graduate school courses as part of our Vesalius Teaching Apprenticeship. The PhD program is designed to produce well-rounded biomedical scientists who expect to include both research and teaching in their career.<\/p>\n<p style=\"text-align: justify;\"><span>&nbsp;<\/span>The <span>PhD curriculum consists of required<\/span> courses<span>, seminars and electives as well as Dissertation Research. <\/span><span>Overview of the 64-credit PhD program:<\/span><\/p>\n<ul class=\"text\" style=\"text-align: left;\">\n<li>Required Courses: 30-32 credits<\/li>\n<li>Electives and\/or Research: 32-34 credits<\/li>\n<li>Optional: Vesalius Module<\/li>\n<\/ul>\n<p style=\"text-align: justify;\">To see more details on the courses we offer, please visit the <a title=\"Course Descriptions\" href=\"https:\/\/www.bumc.bu.edu\/anatneuro\/course-descriptions\/\">Course Description <\/a>page.<\/p>\n<h2 style=\"text-align: left;\"><span style=\"font-size: x-large;\"><span class=\"boldBlue\" style=\"color: #00004f;\">Learning Outcomes<\/span><\/span><\/h2>\n<p style=\"text-align: justify;\">The goals of the Doctor of Anatomy &#038; Neurobiology Program are to provide trainees with the knowledge base and skills to become proficient in basic biomedical research and to develop outstanding pedagogic skills. Successful completion of these goals enables our students to pursue further studies in the biomedical sciences including teaching, research, and health care. At the conclusion of the program students will be able to:<\/p>\n<ul class=\"text\" style=\"text-align: left;\">\n<li>Demonstrate an understanding of Anatomy, Neuroscience, Statistics, and other graduate-level elective anatomical and neuroscience topics through successful completion of our core curriculum.<\/li>\n<li>Demonstrate proficiency in pedagogical skills in the context of the Teaching in Biomedical Sciences course and through teaching fellowships in advanced biomedical courses (as listed above).<\/li>\n<li>Demonstrate proficiency in scientific data acquisition and analysis through mentored research in one of the department\u2019s research labs.<\/li>\n<li>Describe and interpret scientific findings of their laboratory research study through the development of a written, publication-quality thesis.<\/li>\n<li>Demonstrate an understanding of the professional skills (including an ability to read and interpret scientific literature) required of biomedical scientists and educators.<\/li>\n<\/ul>\n<\/div>\n<h2 style=\"text-align: center;\"><span style=\"font-size: x-large;\"><span class=\"boldBlue\" style=\"color: #00004f;\">Required Courses:<\/span><\/span><\/h2>\n<\/div>\n<div class=\"table-wrapper\">\n<table class=\"fl-table\">\n<thead>\n<tr>\n<th style=\"width: 70%;\">Course<\/th>\n<th style=\"width: 15%;\">Credits<\/th>\n<th style=\"width: 15%;\">Code<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<td style=\"width: 350px;\"> Exptl. Design &#038; Statistical Methods<br \/>\n<br \/>\nOR Elementary Biostatistics <\/td>\n<td>2cr<\/td>\n<td>AN704<br \/>\nMS700<\/td>\n<\/tr>\n<tr>\n<td style=\"width: 350px;\"> Professional Skills<\/td>\n<td>2cr<\/td>\n<td>AN715<\/td>\n<\/tr>\n<tr>\n<td> Methods in Neuroscience<\/td>\n<td>4cr<\/td>\n<td>AN718<\/td>\n<\/tr>\n<tr>\n<td> Human Gross Anatomy<br \/>\nOR Histology <\/td>\n<td style=\"width: 50px;\">6\/4cr<\/td>\n<td style=\"width: 100px;\">AN719<br \/>\nAN726<\/td>\n<\/tr>\n<tr>\n<td> Fundamentals of Cell &#038; Molecular Neurobiology<\/td>\n<td>4cr<\/td>\n<td>AN777<\/td>\n<\/tr>\n<tr>\n<td> Research Colloquium (Journal Club)* <\/td>\n<td>2cr<\/td>\n<td>AN801<br \/>\nAN802<\/td>\n<\/tr>\n<tr>\n<td> Vesalius 1: Teaching in the Biomedical Sciences<\/td>\n<td>2cr<\/td>\n<td>AN806<\/td>\n<\/tr>\n<tr>\n<td>Vesalius 2: Teaching Apprenticeship (fulfilled by Teaching Fellow assignments)<\/td>\n<td>2cr<\/td>\n<td>AN809<\/td>\n<\/tr>\n<tr>\n<td> Systems Neurobiology <\/td>\n<td>4cr<\/td>\n<td>AN810<\/td>\n<\/tr>\n<tr>\n<td> Scientific Writing <\/td>\n<td>2cr<\/td>\n<td>AN815<\/td>\n<\/tr>\n<tr>\n<td>One Departmental Seminar Course<\/td>\n<td>2+cr<\/td>\n<td>List Below<\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<\/div>\n<div style=\"padding-right: 50px; padding-left: 50px; padding-bottom: 20px;\">\n<h2 style=\"text-align: center;\"><span style=\"font-size: x-large;\"><span class=\"boldBlue\" style=\"color: #00004f;\">Departmental Advanced Seminar Courses:<\/span><\/span><\/h2>\n<\/div>\n<div class=\"table-wrapper\">\n<table class=\"fl-table\">\n<thead>\n<tr>\n<th style=\"width: 70%;\">Course<\/th>\n<th style=\"width: 15%;\">Credits<\/th>\n<th style=\"width: 15%;\">Code<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<td style=\"width: 350px;\">Neurobiology of Learning and Memory (even years)<\/td>\n<td style=\"width: 50px;\">2cr<\/td>\n<td style=\"width: 100px;\">AN702<\/td>\n<\/tr>\n<tr>\n<td>Neurobiology of Aging (odd years)<\/td>\n<td>2cr<\/td>\n<td>AN707<\/td>\n<\/tr>\n<tr>\n<td>Neurobiology of Visual System (on demand*)<\/td>\n<td>2cr<\/td>\n<td>AN807<\/td>\n<\/tr>\n<tr>\n<td>Dynamic Modeling (on demand*)<\/td>\n<td>2cr<\/td>\n<td>AN820<\/td>\n<\/tr>\n<tr>\n<td>* on demand = 4 or more students<\/td>\n<td><\/td>\n<td><\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<\/div>\n<div style=\"padding-right: 50px; padding-left: 50px; padding-bottom: 20px;\">\n<h2 style=\"text-align: center;\"><span style=\"font-size: x-large;\"><span class=\"boldBlue\" style=\"color: #00004f;\">Department Elective Courses include:<\/span><\/span><\/h2>\n<\/div>\n<div class=\"table-wrapper\">\n<table class=\"fl-table\">\n<thead>\n<tr>\n<th style=\"width: 70%;\">Course<\/th>\n<th style=\"width: 15%;\">Credits<\/th>\n<th style=\"width: 15%;\">Code<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<td> Advanced Clinical Anatomy <\/td>\n<td>2cr<\/td>\n<td>AN708<\/td>\n<\/tr>\n<tr>\n<td> Developmental Cognitive Neuroscience<\/td>\n<td>4cr<\/td>\n<td>AN716<\/td>\n<\/tr>\n<tr>\n<td> Cellular Organization of Tissues <\/td>\n<td>4cr<\/td>\n<td>AN722<\/td>\n<\/tr>\n<tr>\n<td> Advanced Neuroanatomy (even years)\t<\/td>\n<td>2\/4cr<\/td>\n<td>AN724<\/td>\n<\/tr>\n<tr>\n<td> Neuroanatomy Through Clinical Cases <\/td>\n<td>4cr<\/td>\n<td>AN725<\/td>\n<\/tr>\n<tr>\n<td> Vesalius III: Teaching Practicum <\/td>\n<td>2cr<\/td>\n<td>AN805<\/td>\n<\/tr>\n<tr>\n<td> Cognitive Neuroscience <\/td>\n<td>4cr<\/td>\n<td>AN811<\/td>\n<\/tr>\n<tr>\n<td> Human Anatomy and Osteology <\/td>\n<td>4cr<\/td>\n<td>FA712<\/td>\n<\/tr>\n<tr>\n<td> Advanced Human Osteology\t<\/td>\n<td>4cr<\/td>\n<td>FA806<\/td>\n<\/tr>\n<tr>\n<td> Forensic Biology\t<\/td>\n<td>3cr<\/td>\n<td>FS702<\/td>\n<\/tr>\n<tr>\n<td> Forensic Pathology <\/td>\n<td>3cr<\/td>\n<td>FS712<\/td>\n<\/tr>\n<tr>\n<td> Molecular Biology of Forensic DNA <\/td>\n<td>3cr<\/td>\n<td>FS720<\/td>\n<\/tr>\n<tr>\n<td> Biomedical Imaging Foundations <\/td>\n<td>4cr<\/td>\n<td>IM600<\/td>\n<\/tr>\n<tr>\n<td> Methods of Functional Imaging of the Brain<\/td>\n<td>2cr<\/td>\n<td>IM630<\/td>\n<\/tr>\n<tr>\n<td> Molecular Basis of Neurologic Disease<\/td>\n<td>2cr<\/td>\n<td>MS783<\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<\/div>\n<div style=\"padding-right: 50px; padding-left: 50px; padding-bottom: 20px; margin-bottom: 300px; \">\n<h2 style=\"text-align: left;\"><span style=\"font-size: x-large;\"><span class=\"boldBlue\" style=\"color: #00004f;\">Requirements for the PhD Vesalius Module:<\/span><\/span><\/h2>\n<p style=\"text-align: justify;\">Students are required to have taken at least two of the following prerequisite courses to pursue the Vesalius Module:<\/p>\n<p style=\"text-align: left;\">The Module consists of three courses\/components, two of which (Vesalius 1 &#038; 2) are included in the list of required courses for all PhD students. The program begins with a course on the development of teaching skills in the biomedical sciences. This is followed by a teaching apprenticeship (160 hours service as a Teaching Fellow in one or more of the medical or graduate required courses, above), and concludes with a Mentored Teaching Project that involves the development of a didactic lesson or exercise under the direct mentorship of experienced and award-winning faculty.<\/p>\n<div class=\"col-md-6\">\n<h2 style=\"text-align: left; margin: auto;\"><span style=\"font-size: x-large;\"><span class=\"boldBlue\" style=\"color: #00004f;\">Vesalius Module Components:<\/span><\/span><\/h2>\n<div class=\"table-wrapper2\">\n<table class=\"fl-table\">\n<thead>\n<tr>\n<th style=\"width: 70%;\">Course<\/th>\n<th style=\"width: 15%;\">Credits<\/th>\n<th style=\"width: 15%;\">Code<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<td style=\"width: 350px;\">Vesalius 1: Teaching in the Biomedical Sciences<\/td>\n<td style=\"width: 50px;\">2cr<\/td>\n<td style=\"width: 100px;\">AN806<\/td>\n<\/tr>\n<tr>\n<td>Vesalius 2: Teaching Apprenticeship<\/td>\n<td>2cr<\/td>\n<td>AN809<\/td>\n<\/tr>\n<tr>\n<td>Vesalius 3: Mentored Teaching Project (Practicum)<\/td>\n<td>2cr<\/td>\n<td>AN805<\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<\/div>\n<\/div>\n<div class=\"col-md-6\">\n<h2 style=\"text-align: left; margin: auto;\"><span style=\"font-size: x-large;\"><span class=\"boldBlue\" style=\"color: #00004f;\">Vesalius Module Prerequisites (choose 2):<\/span><\/span><\/h2>\n<div class=\"table-wrapper2\">\n<table class=\"fl-table\">\n<thead>\n<tr>\n<th style=\"width: 70%;\">Course<\/th>\n<th style=\"width: 15%;\">Credits<\/th>\n<th style=\"width: 15%;\">Code<\/th>\n<\/tr>\n<\/thead>\n<tbody>\n<tr>\n<td style=\"width: 350px;\">\nHuman Gross Anatomy<\/td>\n<td style=\"width: 50px;\">6cr<\/td>\n<td style=\"width: 100px;\">AN719<\/td>\n<\/tr>\n<tr>\n<td>Graduate Histology<\/td>\n<td>4cr<\/td>\n<td>AN726<\/td>\n<\/tr>\n<tr>\n<td>Systems Neurobiology<\/td>\n<td>4cr<\/td>\n<td>AN810<\/td>\n<\/tr>\n<tr>\n<td>Cognitive Neuroscience<\/td>\n<td>4cr<\/td>\n<td>AN811<\/td>\n<\/tr>\n<\/tbody>\n<\/table>\n<\/div>\n<\/div>\n<\/div>\n<p><!-- ___________________Timeline___________________ --><\/p>\n<section id=timeline>\n<div style=\"padding-top: 0px; padding-right: 50px; padding-left: 50px; padding-bottom: 20px;\">\n<h2 style=\"text-align: center;\"><span style=\"font-size: x-large;\"><span class=\"boldBlue\" style=\"color: #00004f;\">Example of a typical curriculum for PhD students (see required courses and options above):<\/span><\/span><\/h2>\n<\/div>\n<div class=\"demo-card-wrapper\">\n<div class=\"demo-card demo-card--step1\">\n<div class=\"head\">\n<div class=\"number-box\">\n\t\t\t\t\t<span>01<\/span>\n\t\t\t\t<\/div>\n<h2 style=\"color: #ffffff;\"><span class=\"small\">Fall-Spring<\/span> Year 1<\/h2>\n<\/p><\/div>\n<div class=\"body\">\n<p class=\"go-left\"><strong>FALL:<\/strong><\/p>\n<p class=\"go-left\">Systems Neurobiology<br \/>\n4 cr&nbsp;&nbsp;&nbsp; AN 810<\/p>\n<p class=\"go-left\">Embryology<br \/>\n2 cr&nbsp;&nbsp;&nbsp; AN 727<\/p>\n<p class=\"go-left\">Human Gross Anatomy<br \/>\n6 cr&nbsp;&nbsp;&nbsp; AN 719<\/p>\n<p class=\"go-left\"><strong>SPRING:<\/strong><\/p>\n<p class=\"go-left\">Professional Skills<br \/>\n2 cr&nbsp;&nbsp;&nbsp; AN 715<\/p>\n<p class=\"go-left\">Vesalius 1: Teaching in the Biomedical Sciences<br \/>\n2 cr&nbsp;&nbsp;&nbsp; AN 806<\/p>\n<p class=\"go-left\">Methods in Neuroscience<br \/>\n4 cr&nbsp;&nbsp;&nbsp; AN 718<\/p>\n<p class=\"go-left\">Research Colloquium (Journal Club)<br \/>\n2 cr&nbsp;&nbsp;&nbsp; AN 802<\/p>\n<p class=\"go-left\">Elective(s) (see list above)<br \/>\n2 cr&nbsp;&nbsp;&nbsp; <\/p>\n<p class=\"go-left\"><em>PhD students typically rotate through 2-3 labs during the Spring semester. This enables them (and faculty research mentors) to find a lab in which dissertation research will be performed.<\/em><\/p>\n<\/div>\n<\/div>\n<div class=\"demo-card demo-card--step2\">\n<div class=\"head\">\n<div class=\"number-box\">\n\t\t\t\t\t<span>02<\/span>\n\t\t\t\t<\/div>\n<h2><span class=\"small\">Fall-Spring<\/span> Year 2<\/h2>\n<\/p><\/div>\n<div class=\"body\">\n<p><strong>FALL:<\/strong><\/p>\n<p>Exptl. Design &amp; Statistical Methods or Elementary Biostats.<br \/>\n2 cr&nbsp;&nbsp;&nbsp; AN 704\/MS 700\/MS 750\/BS 704<\/p>\n<p>Fundamentals of Cell &amp; Molecular Neurobiology<br \/>\n4 cr&nbsp;&nbsp;&nbsp; AN 777<\/p>\n<p>Scientific Writing<br \/>\n2 cr&nbsp;&nbsp;&nbsp; AN 815<\/p>\n<p><strong>SPRING:<\/strong><strong> (<\/strong>Note this is when most students take their PhD qualifying exam)<\/p>\n<p>Advanced Seminar (1 required, see list above)<br \/>\n2 cr&nbsp;&nbsp;&nbsp; (see list above)<\/p>\n<p>Elective(s)<br \/>\nVariable<\/p>\n<\/div>\n<\/div>\n<div style=\"height: 475px;\" class=\"demo-card demo-card--step3\">\n<div class=\"head\">\n<div class=\"number-box\">\n\t\t\t<span>03<\/span>\n\t\t<\/div>\n<h2 style=\"color: #ffffff;\"><span class=\"small\">Fall-Spring<\/span> Year 3<\/h2>\n<\/p><\/div>\n<div  style=\"height: 400px;\" class=\"body\">\n<p><strong>FALL:<\/strong><\/p>\n<p>Research<br \/>\nVariable<\/p>\n<p>Elective(s) (see list above)<br \/>\nVariable<\/p>\n<p>Vesalius 2: Teaching Apprenticeship&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; (continues to 160 hrs)<br \/>\n6 Cr&nbsp;&nbsp;&nbsp; AN 805<\/p>\n<p><strong>SPRING:<\/strong><\/p>\n<p>Research<br \/>\nVariable<\/p>\n<p>Elective(s) (see list above)<br \/>\nVariable<\/p>\n<p>Advanced Seminar (see list above)<br \/>\nVariable<\/p>\n<\/div>\n<\/div>\n<div  style=\"height: 275px;\" class=\"demo-card demo-card--step4\">\n<div class=\"head\">\n<div class=\"number-box\">\n\t\t\t\t\t<span>04<\/span>\n\t\t\t\t<\/div>\n<h2><span class=\"small\">Fall-Spring<\/span> Year 4<\/h2>\n<\/p><\/div>\n<div style=\"height: 200px;\" class=\"body\">\n<p><strong>SPRING-FALL:<\/strong><\/p>\n<p>Research and Dissertation Preparation<br \/>\nVariable<\/p>\n<\/div>\n<\/div>\n<div style=\"height: 275px;\" class=\"demo-card demo-card--step5\">\n<div class=\"head\">\n<div class=\"number-box\">\n\t\t\t\t\t<span>05<\/span>\n\t\t\t\t<\/div>\n<h2 style=\"color: #ffffff;\"><span 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